Valid point. primarily vegetarian (55-85% plant matter depending on time of year).Nope. Even the paper you linked disagrees with this. They are predominantly herbivorous but their diet varies with the seasons and also their location. When they're available, insects, insect larvae, worms and crustaceans are also eaten.
This can easily be verified by the examined stomach contents given on fishbase and nearly all studies to date. The paper you quote, while recognising discus as predominantly herbivorous, also states...
"a large proportion of discus exhibited only partially filled stomachs at low water, and... invertebrates comprised a greater fraction of their diet. During the high water period discus were usually found in open, well lit flooded forests where periphyton grows most profusely. At this time of the year, invertebrates constituted a surprisingly small volumetric component of discus diet, although they may represent a vital source of protein."
By definition, discus are not "vegetarian"".
They do generally get fed too much protein in captivity though, with beefheart being the most ridiculous choice, and presumably the most damaging to their long-term health.
Most fish can't digest mammal meat very well, even mammals don't digest mammal meat that well when it's raw.