Problem with a lot of species is that several were long be seen as the same species and got seperated species later.
C.aeneus - C.venezuelanus - C.schultzei were earlier considered as C.aeneus. When they mated it wasn't considered crossbreeding.
That's the way green Aeneus were bred (there is a natural type also btw).
The seperate species C.macrosteus (huge Corys) is now C.aeneus again.
Same with C.paleatus and C.longipinnis.
Now the offspring of these are considered crrossbreds / hybrids. Earlier they weren't
So some fish got crossbred without the intention to do so.
I concur with these statements and suspected this myself over the years and this is a good explanation of this imho.
Nicely done.