Predators.
Birds.
As aquarists, we try to arrange our tanks to minimize stress, but the stress faced by different Cory species across their enormous geographic range will differ. Is the species adapted to rainforest, under the canopy streams where their instincts are tuned to shade and shadows? Are they from more open habitats? Do they respond to differently coloured substrates, in relation to their camouflage? Is light and fast water their cover? How large are the shoals they move in, in nature?
We can influence how instinct expresses itself, but the most important thing to remember is these are not aquarium fish. They are species evolved in a tough habitat, some of whom happened to be caught and put into the aquarium trade. Instinct does not vanish in a few generations. And since the very basis of what a species is involves difference, it's going to show in behaviour as well as size and colouration.
I was thinking the same thing, didn't know how to express though, and I wouldn't have been able to do it so eloquently and concisely!
From my own experience with cories, I've found bronzes to be the lovable goofballs dopes of the cory world. The labs/goldies o the cory groups! Lovable, but a bit dim...
They've been tank raised for so long, and they're just known as being less easily spooked. While my sterbai are more alert to potential dangers and tend to hide away more. My pygmy cories were much more inclined to hide away when I only had a group of seven of them, and they showed much more confidence and different behaviours, like swimming around in large groups, like schoaling, once they were in a group of 20 or more. I did see that they still spooked - there were more eyes watching out for danger for one thing, and once one spooked, it tends to spook most-all of them. But being in a larger group also meant they recovered from the spooking much faster, and came back out into the open again sooner.
I was watching the mink man the other day, who uses mink and dogs to help eradicate pest rats around farms and things. He showed that muskrats were so absolutely terrified of mink that they'd run and even hide behind or try to fight a human, rather than risk getting trapped in their burrow by the mink. When he sends a mink into a hidey hole to catch rats, the other rats in there, even while a mink is killing another rat, often stay hidden in there with the mink, rather than try to run and face the dogs and humans outside the burrow. Some run, but many don't. Makes sense. Mink naturally predate on muskrats, share the same habitats, muskrats have evolved knowing that mink are a huge danger to them. Rats that are found around human areas don't really encounter mink, but humans and dogs are a huge danger for them. So they'd rather stay where the mink is, even seeing the danger happening, than run into the things they've evolved to be afraid of.
Different environments apply different pressures to different species, and it's bound to show in more than just their physiology.